1. Division of Bioresources, Hokkaido University International Institute for Zoonosis Control, Sapporo 001-0020, Japan
2. Department of Veterinary Public Health, Faculty of Veterinary Medicine, Kasetsart University, Bangkok 73140, Thailand
3. Hokkaido University Institute for Vaccine Research & Development, Hokkaido University, Sapporo 001-0020, Japan
4. International Collaboration Unit, Hokkaido University International Institute for Zoonosis Control, Sapporo 001-0020, Japan