1. Division of Plant Sciences, School of Life Sciences, University of Dundee, Dow Street, Dundee DD1 5EH, Scotland, UK
2. Information and Computational Sciences, The James Hutton Institute, Invergowrie, Dundee DD2 5DA, Scotland, UK
3. Cell and Molecular Sciences, The James Hutton Institute, Invergowrie, Dundee DD2 5DA, Scotland, UK