1. Institute for Molecular and Cellular Biology, Osaka University, and CREST, Japan Science and Technology Corporation, 1-3 Yamada-oka, Suita, Osaka 565-0871, Japan, The Graduate School of Pharmaceutical Sciences, Osaka University, 1-6 Yamada-oka, Suita, Osaka 565-0872, Japan, Biomolecular Engineering Research Institute, 6-2-3 Furuedai, Suita, Osaka 565-0874, Japan, and RIKEN (The Institute of Physical and Chemical Research), Wako-shi, Saitama 351-0198, Japan