1. RIKEN Center for Biosystems Dynamics Research, Suita, Osaka 565-0874, Japan
2. Graduate School of Frontier Biosciences, Osaka University, Suita, Osaka 565-0871, Japan
3. Research Headquarters, Taisho Pharmaceutical Co., LTD., Saitama-shi, Saitama 331-9530, Japan
4. Research Institute for Radiation Biology and Medicine, Hiroshima University, Hiroshima City, Hiroshima 734-8553, Japan