1. Department of Pharmacology, Keio University School of Medicine, 35 Shinanomachi, Shinjuku, Tokyo 160-8582, Japan
2. Graduate School of Environment and Information Sciences, Yokohama National University, Kanagawa 240-8501, Japan
3. Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Agency (JST), Kawaguchi 332-0012, Saitama, Japan