1. CAS Key Laboratory of Biobased Materials, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, No. 189 Songling Road, Qingdao 266101, China
2. University of Chinese Academy of Sciences, No. 19(A) Yuquan Road, Beijing 100049, China
3. State Key Laboratory of Microbial Technology, Shandong University, No. 72 Binhai Road, Qingdao 266237, China
4. Qingdao Youheng Biotechnology Co., Ltd., No. 130 Jiushui East Road, Qingdao 266199, China