1. Harvard−MIT Division of Health Sciences and Technology, Massachusetts Institute of Technology, Cambridge, Massachusetts 02139, United States
2. Medical Scientist Training Program, Harvard Medical School, Boston, Massachusetts 02115, United States
3. Broad Institute of Harvard and MIT, Cambridge, Massachusetts 02142, United States
4. Department of Medicine, Brigham and Women’s Hospital, Boston, Massachusetts 02115, United States
5. Electrical Engineering and Computer Science, David H. Koch Institute for Integrative Cancer Research, Massachusetts Institute of Technology, Cambridge, Massachusetts 02139, United States
6. Howard Hughes Medical Institute, Chevy Chase, Maryland 20815, United States