1. Flavor Science Innovation Center, Technology Research Center, Wuliangye Yibin Co., Ltd., Yibin, Sichuan 644007, China
2. Solid-State Fermentation Resource Utilization Key Laboratory of Sichuan Province, Yibin, Sichuan 644007, China
3. Key Laboratory of Wuliangye-Flavor Liquor Solid-State Fermentation, China National Light Industry, Yibin, Sichuan 644007, China
4. Department of Food Science and Technology, Oregon State University, Corvallis, Oregon 97331, United States