1. College of Natural Resources and Environment, South China Agricultural University, Guangzhou 510642, China
2. Laboratory of Ecology, Faculty of Sciences and Technology, Cheikh Anta University of Dakar Dakar 50005, Senegal
3. Guangdong Provincial Key Laboratory of Utilization and Conservation of Food and Medicinal Resources in Northern Region, Shaoguan 512005, China
4. School of Biology and Agriculture, Shaoguan University, Shaoguan 512005, China