1. Guangxi Laboratory on the Study of Coral Reefs in the South China Sea, Coral Reef Research Center of China, School of Marine Sciences, Guangxi University, Nanning 530004, China
2. Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519080, China
3. Guangzhou Institute of Geochemistry, Chinese Academy of Sciences, Guangzhou 510640, China