1. Graduate School of Human Development and Environment, Kobe University, 3-11 Tsurukabuto, Nada-ku, Kobe City, Hyogo 657-8501, Japan
2. Research Fellow of Japan Society for the Promotion of Science, 5-3-1 Kojimachi, Chiyoda-ku, Tokyo, 102-0083, Japan
3. Faculty of Human Development, Kobe University, 3-11 Tsurukabuto, Nada-ku, Kobe City, Hyogo 657-8501, Japan
4. Maizuru Fisheries Research Station, Field Science Education and Research Center, Kyoto University, Maizuru, Kyoto 625-0086, Japan