1. Department of Biotechnology, Graduate School of Engineering, Nagoya University, Furo-cho, Chikusa-ku, Nagoya, Aichi 464-8603, Japan
2. Department of Biomolecular Engineering, Graduate School of Engineering, Nagoya University, Furo-cho, Chikusa-ku, Nagoya, Aichi 464-8603, Japan
3. Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki 305-0006, Japan
4. Faculty of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki 305-0006, Japan
5. Microbiology Research Center for Sustainability, University of Tsukuba, Tsukuba, Ibaraki 305-8572, Japan