1. National Oceanic and Atmospheric Administration, Northwest Fisheries Science Center, Seattle, Washington 98112, United States
2. National Oceanic and Atmospheric Administration, Saltwater, Inc., under Contract to Northwest Fisheries Science Center, Seattle, Washington 98112, United States
3. Marrowstone Marine Field Station, US Geological Survey, Western Fisheries Research Center, Nordland, Washington 98358-9633, United States
4. Institute of Marine Research, Bergen, Nordnes 5817, Norway