1. Center of Wastewater Resource Recovery, College of Resources and Environment, Fujian Agriculture and Forestry University, Fuzhou 350002, China
2. Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao 266101, China
3. Department of Biotechnology, Delft University of Technology, Julianalaan 67, Delft 2628 BC, The Netherlands