1. Key Laboratory of Tropical Marine Bio-resources and Ecology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China
2. Innovation Academy of South China Sea Ecology and Environmental Engineering, Chinese Academy of Sciences, Guangzhou 510301, China
3. School of Life and Environmental Sciences, Centre for Integrative Ecology, Deakin University, Burwood, Victoria 3125, Australia
4. Department of Environmental Sciences, University of Virginia, Charlottesville, Virginia 22911, United States