1. Karen Foster Environmental Research, Peterborough, ON K9J 8L2, Canada
2. Applications of Modelling & Quantitative Methods (AMOD), Trent University, Peterborough, ON K9L 0G2, Canada
3. Environment and Climate Change Canada, National Wildlife Research Centre, Carleton University, Ottawa, ON K1A 0H3, Canada
4. Biology Department, Acadia University, Wolfville, NS B4P 2R6, Canada