1. School of Biosciences, University of Birmingham, Birmingham B15 2TT, United Kingdom
2. School of Marine Science and Technology, Newcastle University, Newcastle upon Tyne NE1 7RU, United Kingdom
3. TNO Science and Industry, Bevesierweg MML (Harssens), Den Helder, The Netherlands
4. Division of Mechanical Engineering, University of Dundee, Dundee, DD1 4HN, United Kingdom