1. The Edison Family Center for Genome Sciences & Systems Biology, Washington University School of Medicine, St. Louis, Missouri 63110, United States
2. Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri 63110, United States
3. Department of Biomedical Engineering, Washington University in St. Louis, St. Louis, Missouri 63130, United States
4. Department of Molecular Microbiology, Washington University School of Medicine, St. Louis, Missouri 63110, United States