1. Department of Engineering Science, University of Oxford, Parks Road, Oxford OX1 3PJ, United Kingdom
2. Hangzhou Innovation Center and College of Chemical & Biological Engineering, Zhejiang University, Hangzhou 311200, China
3. School of Biological Sciences, University of Edinburgh, G20 Roger Land Building, The Kingʼs Buildings, Edinburgh EH9 3FF, United Kingdom
4. ZJU-UoE Joint Research Centre for Engineering Biology, Zhejiang University, Haining 314400, China