1. Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences, Tianjin 300308, China
2. University of Chinese Academy of Science, Beijing 100049, China
3. Key Laboratory of Engineering Biology for Low-Carbon Manufacturing, Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences, Tianjin 300308, China
4. National Center of Technology Innovation for Synthetic Biology, Tianjin 300308, China
5. College of Biotechnology, Tianjin University of Science & Technology, Tianjin 300457, China