1. National Engineering Research Center for Breeding Swine Industry, South China Agricultural University, Guangzhou 510642, China
2. Department of Animal Genetics, Breeding and Reproduction, College of Animal Science, South China Agricultural University, Guangzhou 510642, China
3. Guangdong Provincial Key Laboratory of Agro-Animal Genomics and Molecular Breeding, South China Agricultural University, Guangzhou 510642, China
4. State Key Laboratory Swine and Poultry Breeding Industry, South China Agricultural University, Guangzhou 510642, China
5. National and local joint Engineering Research Center for Livestock and Poultry Breeding Industry, South China Agricultural University, Guangzhou 510642, China