1. Jiangsu Key Laboratory for the Research and Utilization of Plant Resources, Institute of Botany, Jiangsu Province and Chinese Academy of Sciences (Nanjing Botanical Garden Mem. Sun Yat-Sen), Nanjing 210014, China
2. The New Zealand Institute for Plant and Food Research Ltd (PFR), Private Bag, 92169 Auckland, New Zealand
3. College of Horticulture, Nanjing Agricultural University, Nanjing 210095, China
4. State Key Laboratory of Crop Genetics & Germplasm Enhancement and Utilization, Nanjing Agricultural University, Nanjing 210095, China