1. Crop Research Institute, Hunan Academy of Agricultural Sciences, Changsha 410125, People’s Republic of China
2. Yuelushan Laboratory, Changsha 410128, People’s Republic of China
3. School of Life Sciences, Chinese University of Hong Kong, Hong Kong 999077, People’s Republic of China
4. College of Agronomy, Hunan Agricultural University, Changsha 410128, People’s Republic of China
5. Hunan Key Laboratory of Economic Crops Genetic Improvement and Integrated Utilization, School of Life Science, Hunan University of Science and Technology, Xiangtan 411201, People’s Republic of China
6. Key Laboratory of Hunan Province on Crop Epigenetic Regulation and Development, Hunan Agricultural University, Changsha 410128, People’s Republic of China
7. College of Bioscience and Biotechnology, Hunan Agricultural University, Changsha, Hunan 410128, China