1. Plant Breeding, Wageningen University and Research, Wageningen 6708 PB, The Netherlands
2. State Key Laboratory of Vegetable Biobreeding, Key Laboratory of Biology and Genetic Improvement of Horticultural Crops of the Ministry of Agriculture and Rural Affairs, Sino-Dutch Joint Laboratory of Horticultural Genomics, Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Beijing 100081, China
3. Bioscience, Wageningen University and Research, Wageningen 6708 PB, The Netherlands