1. State Key Laboratory of Microbial Technology, Shandong University, Qingdao266237, China
2. School of Life Sciences, University of Warwick, CoventryCV4 7AL, United Kingdom
3. College of Marine Life Sciences, and Frontiers Science Center for Deep Ocean Multispheres and Earth System, Ocean University of China, Qingdao266003, China
4. Laboratory for Marine Biology and Biotechnology, Pilot National Laboratory for Marine Science and Technology, Qingdao266237, China
5. Marine Biotechnology Research Center, State Key Laboratory of Microbial Technology, Shandong University, Qingdao266237, China