1. Department of Computer Science, University of Bristol, Woodland Road, Bristol BS8 1UB, UK
2. Department of Ecology and Evolutionary Biology, University of Arizona, PO Box 210088, Tucson, AZ 85721, USA
3. Department of Mathematics, VU University Amsterdam, De Boelelaan 1081a, 1081 HV Amsterdam, The Netherlands
4. School of Biological Sciences, University of Bristol, Woodland Road, Bristol BS8 1UG, UK