1. Insititute of Evolution and Marine Biodiversity, Ocean University of China, Qingdao 266003, People's Republic of China
2. Department of Biological Sciences, Smith College, Northampton, MA 01063, USA
3. Key Laboratory of Mariculture (Ocean University of China), Ministry of Education, Qingdao 266003, People's Republic of China
4. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266237, People's Republic of China