1. School of Biosciences, Cardiff University, Cardiff CF10 3AX, UK
2. Department of Biodiversity and Molecular Ecology, Research and Innovation Centre, Fondazione Edmund Mach, S. Michele all'Adige, Trentino 38010, Italy
3. Division of Animal, Food and Health Sciences, CSIRO, Armidale, New South Wales 2350, Australia
4. School of Biological Sciences, University of Bristol, Bristol BS8 1TQ, UK