1. Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow G61 1QH, UK
2. London School of Hygiene and Tropical Medicine, Keppel Street, London WC1E 7HT, UK
3. Global and Tropical Health Division, Menzies School of Health Research and Charles Darwin University, Darwin, Northern Territory 0810, Australia
4. Gleneagles Kota Kinabalu Hospital, 88100, Kota Kinabalu, Sabah, Malaysia
5. Infectious Diseases Society, Sabah-Menzies School of Health Research Clinical Research Unit, Kota Kinabalu 88560, Sabah, Malaysia
6. Royal (Dick) School of Veterinary Studies and Roslin Institute, University of Edinburgh, Easter Bush Campus, Roslin, Midlothian EH25 9RG, UK