1. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, People's Republic of China
2. Department of Earth and Environmental Sciences, Palaeontology and Geobiology, Ludwig-Maximilians-Universität München, Richard-Wagner-Strasse 10, 80333 München, Germany
3. School of Geography, Queen Mary University of London, Mile End Road, London E1 4NS, UK
4. University of Chinese Academy of Sciences, Beijing 100049, People's Republic of China