1. State Key Laboratory of Grassland Agro-ecosystem, College of Ecology, Lanzhou University, Lanzhou, People's Republic of China
2. Department of Anthropology, University College London, London, UK
3. BirthRites Independent Max Planck Research Group, Max Planck Institute for Evolutionary Anthropology, Leipzig, Germany
4. Department of Population Health, London School of Hygiene and Tropical Medicine, London, UK