1. Institute of Virology and Environmental Microbiology, Mansfield Road, Oxford OX1 3SR, UK
2. Biodiversity and Ecology Division, School of Biological Sciences, University of Southampton, Southampton SO17 1BJ, UK
3. Department of Biological and Molecular Sciences, University of Stirling, Stirling FK9 4LA, UK
4. School of Biological and Earth Sciences, Liverpool John Moores University, Byrom Street, Liverpool L3 3AF, UK