1. Centre for Comparative Infectious Diseases, University of Liverpool, Biosciences Building, Liverpool L69 7ZB, UK
2. School of Biological Sciences, University of Liverpool, Biosciences Building, Liverpool L69 72B, UK
3. Aberdeen Population Ecology Research Unit (APERU), Department of Zoology, University of Aberdeen, Tillydrone Avenue, Aberdeen AB24 2TZ, UK
4. Centre for Ecological and Evolutionary Synthesis (CEES), Department of Biology, University of Oslo, PO Box 1050 Blindern, 0316 Oslo, Norway
5. Department of Veterinary Pathology, University of Liverpool, Biosciences Building, Liverpool L69 72B, UK
6. APERU, Centre for Ecology and Hydrology, Banchory Research Station, Hill of Brathens, Banchory AB31 4BW, UK
7. National Mycobacteria Reference Laboratory, National Institute of Public Health and the Environment, PO Box 1, 3720 BA Bilthoven, The Netherlands