1. Department of Biology, University of York, Wentworth Way, York YO10 5DD, UK
2. Butterfly Conservation, Manor Yard, East Lulworth, Wareham BH20 5QP, UK
3. AgroEcology Department, Rothamsted Research, Harpenden AL5 2JQ, UK
4. Centre for Ecology and Conservation, and Environment and Sustainability Institute, University of Exeter, Penryn TR10 9EZ, UK
5. British Trust for Ornithology, The Nunnery, Thetford IP24 2PU, UK
6. Conservation Science Group, Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK
7. Centre for Ecology and Hydrology, Wallingford OX10 8BB, UK