1. Global Change and Biodiversity Programme, South African National Biodiversity Institute, Kirstenbosch Research CentrePrivate Bag X7, Claremont 7735, Republic of South Africa
2. Climate Change Vulnerability and Adaptation Programme, NRF/DST Centre of Excellence at the FitzPatrick Institute of African Ornithology, University of Cape TownRondebosch, Cape Town 7700, Republic of South Africa
3. Laboratoire d'Ecologie Alpine, UMR-CNRS 5553, Université Joseph FourierBP 53, 38041 Grenoble Cedex 9, France