1. Department of Biological Sciences, Macquarie UniversitySydney, New South Wales, Australia, 2109
2. MUCEP, Department of Earth and Planetary Sciences, Macquarie UniversitySydney, New South Wales, Australia, 2109
3. Museum VictoriaPO Box 666E, Melbourne, Victoria 3000, Australia
4. UMR 5143 CNRS, Muséum National d'Histoire Naturelle8 rue Buffon, Paris 75005, France
5. Monash UniversityClayton, Victoria 3168, Australia