1. Department of Basic Science, Graduate School of Arts and Sciences, University of Tokyo, 3-8-1 Komaba, Meguro-ku, Tokyo 153-8902, Japan
2. Research Center for Complex Systems Biology, Graduate School of Arts and Sciences, University of Tokyo, 3-8-1 Komaba, Meguro-ku, Tokyo 153-8902, Japan
3. PRESTO, Japan Science and Technology Agency, Kawaguchi-shi, Saitama 332-0012, Japan