1. Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, NJ, USA
2. Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ, USA
3. Princeton School of Public and International Affairs, Princeton University, Princeton, NJ, USA
4. Department of Mathematics and Statistics, University of Victoria, Victoria, British Columbia, Canada
5. Department of Molecular Biology, Princeton University, Princeton, NJ, USA