1. Research Institute for Global Change (RIGC), JAMSTEC , Yokosuka, Kanagawa 237-0061, Japan
2. Sugashima Marine Biological Laboratory, Graduate School of Science, Nagoya University, Sugashima , Toba, Mie 517-0004, Japan
3. Centre for Marine & Coastal Studies, Universiti Sains Malaysia , Penang 11800, Malaysia
4. Graduate School of Sciences and Technology for Innovation, Graduate School of Sciences and Technology for Innovation, Yamaguchi University , Yamaguchi, Yamaguchi 753-8512, Japan
5. Graduate School/Faculty of Science, Hokkaido University , Sapporo 060-0810, Japan