1. Biology Department, University of Wisconsin Eau Claire, 105 Garfield Avenue, Eau Claire, WI 54702, USA
2. Biology Department, Mills College, 5000 MacArthur Boulevard, Oakland, CA 94631, USA
3. Institute of Evolutionary Medicine, University of Zurich, Zurich 8057, Switzerland
4. School of Human Evolution and Social Change, Institute of Human Origins, Arizona State University, Tempe, AZ 85287-2402, USA