1. Mitchell Center for Sustainability Solutions, University of Maine, Orono, ME 04469, USA
2. Department of Biology, Colby College, 4000 Mayflower Hill Drive, Waterville, ME 04901, USA
3. Institute of Evolution, Centre for Ecological Research, Budapest, Hungary
4. Center for the Conceptual Foundations of Science, Parmenides Foundation, Pöcking, Germany
5. Plant Systematics, Ecology and Theoretical Biology, Eötvös University, Budapest, Hungary