1. 2Department of Biological Sciences, Southern Illinois University at Edwardsville, Edwardsville, Illinois 62026 USA;
2. 3Department of Evolution, Ecology and Organismal Biology, The Ohio State University, Columbus, Ohio 43210 USA;
3. 4Department of Biology, Ashland Uiversity, Ashland, Ohio 44805 USA;
4. 5Department of Higher Plant Systematics and Evolution, Institute of Botany, University of Vienna, Rennweg 14, A‐1030 Vienna, Austria;
5. 6Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs, Connecticut 06269 USA; and
6. 7Departamento de Bota´nica, Universidad de Concepcio´n, Concepcio´n, Chile