1. 2Department of Biology, University of New Mexico, Albuquerque, New Mexico 87131 USA
2. 3School of Biological Science, University of New South Wales, Sydney, New South Wales 2052, Australia
3. 4Jepson Herbarium and Department of Integrative Biology, University of California, Berkeley, California 94720 USA
4. 5Department of Evolution, Ecology & Organismal Biology, Ohio State University, 1735 Neil Avenue, Columbus, Ohio 43210–1220 USA