1. Department of Ecology and Evolutionary Biology, and the Natural History Museum and Biodiversity Research Center, University of Kansas; Lawrence, Kansas 66045 U.S.A.
2. Department of Biology, Indiana University of Pennsylvania; Indiana, Pennsylvania 15705 U.S.A.
3. Jardín de Aclimatación de la Orotava, Puerto de la Cruz, Tenerife; Canary Islands Spain