1. 2Department of Organismic and Evolutionary Biology, Harvard University Herbaria, 22 Divinity Avenue, Cambridge, Massachusetts 02138 USA;
2. 3University of Michigan Herbarium, North University Building, Ann Arbor, Michigan 48109–1057 USA;
3. 4Department of Ecology and Evolutionary Biology, Yale University, P.O. Box 208106, New Haven, Connecticut 06520 USA