1. a John Innes Centre , Colney Lane, Norwich, NR47UH, United Kingdom
2. b Laboratoire des Interactions Plantes Microorganismes (LIPM) , UMR CNRS , INRA 2594/441, Chemin de Borderouge, BP 52627, Castanet Tolosan, 31326, France
3. c School of Computing Sciences , University of East Anglia , Norwich, NR4 7TJ, United Kingdom
4. d División de Genética, and Instituto de Bioingenieria , Universidad Miguel Hernández , Campus de Elche, Elche, 02302, Alicante, Spain
5. e John Innes Centre , Colney Lane, Norwich, NR47UH, United Kingdom E-mail: