1. State Key Laboratory of Plateau Ecology and Agriculture College of Agriculture and Animal Husbandry, Qinghai University Xining, Qinghai 810016, China
2. State Key Laboratory of Plateau Ecology and Agriculture College of Agriculture and Animal Husbandry, Qinghai University Xining, Qinghai 810016, China; College of Agriculture and Animal Husbandry, Qinghai University Xining, Qinghai 810016, China