1. Interdisciplinary Graduate School of Science and Engineering, Shimane University, 1060 Nishikawatsu-cho, Matsue 690-8504, Japan (e-mail: )
2. Institute of Environmental Systems Science, Academic Assembly, Shimane University, 1060 Nishikawatsu-cho, Matsue 690-8504, Japan
3. Estuary Research Center, Shimane University, 1060 Nishikawatsu-cho, Matsue 690-8504, Japan
4. National Institute of Polar Research, 10-3 Midoricho, Tachikawa, Tokyo 190-8518, Japan