1. Centre for Sustainable Agriculture, Rural Development and Extension, Faculty of Natural and Agricultural Sciences, University of the Free State, PO Box 339, Bloemfontein 9300, South Africa
2. Department of Geography/Afromontane Research Unit, Faculty of Natural and Agricultural Sciences, University of the Free State, PO Box 339, Phuthaditjhaba 9866, South Africa
3. Department of Zoology and Entomology/Afromontane Research Unit, Faculty of Natural and Agricultural Sciences, University of the Free State, PO Box 339, Phuthaditjhaba 9866, South Africa