1. Dogs Trust, London EC1V 7RQ, UK
2. Institute for Advanced Study, Kyoto University, Kyoto 6068501, Japan
3. Japan Society for the Promotion of Science, Tokyo 1020083, Japan
4. Bristol Veterinary School, University of Bristol, Bristol BS40 5DU, UK
5. Linnaeus Veterinary Limited, Shirley, Solihull B90 4BN, UK